Eumetazoa (from Ancient Greekεὖ (eû)'well'μετά (metá)'after' and ζῷον (zôion)'animal'), also known as Epitheliozoa or Histozoa, is a proposed basalanimalsubkingdom as a sister group of Porifera (sponges).[8][9][10][11][12] The basal eumetazoan clades are the Ctenophora and the ParaHoxozoa. Placozoa is now also seen as a eumetazoan in the ParaHoxozoa. The competing hypothesis is the Myriazoa clade.[13] The subkingdom Parazoa and Agnotozoa are the other taxa, and agnotozoa may be fake or even nonexistent at studies. Parazoa or Agnotozoa are a main sister group to eumetazoans, forming clade Blastozoa/Diploblastozoa. Alternatively,
Parazoa was considered as a sister group to Agnotozoa(now considered polyphyletic).
Several other extinct or obscure life forms, such as Iotuba and Thectardis, appear to have emerged in the group.[14] Characteristics of eumetazoans include true tissues organized into germ layers, the presence of neurons and muscles, and an embryo that goes through a gastrula stage.
Some phylogenists once speculated the sponges and eumetazoans evolved separately from different single-celled organisms, which would have meant that the animal kingdom does not form a clade (a complete grouping of all organisms descended from a common ancestor). However, genetic studies and some morphological characteristics, like the common presence of choanocytes, now unanimously support a common origin.[15]
Traditionally, eumetazoans are a major group of animals in the Five Kingdoms classification of Lynn Margulis and K. V. Schwartz, comprising the Radiata and Bilateria – all animals except the sponges.[16].
^Tang, F.; Bengtson, S.; Wang, Y.; Wang, X.L.; Yin, C.Y. (20 September 2011). "Eoandromeda and the origin of Ctenophora". Evolution & Development. 13 (5): 408–414. doi:10.1111/j.1525-142X.2011.00499.x. PMID23016902. S2CID 28369431.
^Fedonkin, M. A. (1990). Systematic description of Vendian Metazoa. In: Sokolov, B. S. & Iwanowski, A. B. (eds), *The Vendian System: Historical–Geological and Paleontological Foundation*, Vol. 1. Springer, pp. 71–120. Fedonkin describes trilobozoans as coelenterate-like organisms, implying eumetazoan affinities.
^Lankester, Ray (1877). "Notes on the Embryology and classification of the Animal kingdom: comprising a revision of speculations relative to the origin and significance of the germ-layers". Quarterly Journal of Microscopical Science (N.S.), No. 68: 399–454.
^Beklemishev, V. L. The basis of the comparative anatomy of the invertebrates [Основы сравнительной анатомии беспозвоночных]. 1st ed., 1944; 2nd ed., 1950; 3rd ed. (2 vols.), 1964. English translation, 1969, [1]. Akademia Nauk, Moscow, Leningrad.
^Ulrich, W. (1950). "Begriff und Einteilung der Protozoen". In Grüneberg, H. (ed.). Moderne Biologie. Festschrift zum 60. Geburtstag von Hans Nachtsheim (in German). Berlin: Peters. pp. 241–250.